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Unassembled sequences in the Global Ocean Sampling metagenomic data independently verified this global scale geographical signal in some Roseobacter genes. The primary findings emerging from this comparative genome analysis are that members of the lineage cannot be easily collapsed into just a few ecologically differentiated clusters (that is, there are almost as many clusters as isolates); the strongest framework for predicting genome content is trophic strategy, but no single framework gives robust predictions; and previously unknown homologs to genes for H2 oxidation, proteorhodopsin based phototrophy, xanthorhodpsin based phototrophy, and CO2 fixation by Form IC ribulose 1,5 bisphosphate carboxylase (RuBisCO) expand the possible mechanisms for energy and carbon acquisition in this remarkably versatile bacterial lineage.Keywords: genome; marine; phylogeny; RoseobacterTop of pageIntroductionSince the discovery of their abundance in marine bacterioplankton communities two decades ago (Gonz and Moran, 1997), members of the marine Roseobacter lineage have emerged as important model organisms for marine microbial ecology. The group spans multiple described genera (at least 45), encompasses a comparatively large sequence variation among 16S rRNA genes (up to 11 and has a poorly resolved within taxon phylogeny (Buchan et al., 2005; Wagner D and Biebl, 2006; Brinkhoff et al., 2008).

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